ENDOMEMBRANE SYSTEM
- It is a grouping of some membrane bound organelles which function in close coordination with each other , i.e. , endoplasmic reticulum , Golgi complex , lysosomes and vacuoles .
- ER was discovered by Porter and Thompson in 1945.
- It is the largest of the internal membranes .
- ER is an interconnected system of membrane lined channels that run through cytoplasm .
- It is complicated organelle that exists in three forms cisternae , vesicles and tubules.
- It is of two types smooth endoplasmic reticulum ( SER , without ribosomes ) and rough endoplasmic reticulum ( RER , with ribosomes ) .
- RER provides surface for protein synthesis .
- The proteins in ER lumen are processed and are packed in membrane bound vesicles for storage .
- SER is involved in the synthesis of lipids and other biosynthetic activities .
Golgi Complex
- At various locations within the endomembrane system , flattened stacks of membranes occur which are often interconnected with one another .
- There are known as Golgi bodies and collectively referred to as the Golgi apparatus .
- The Golgi apparatus was discovered by Camillo Golgi in 1898 and was also named after him .
- It is a parallel arrangement and interconnected system of a cluster of smooth membranous disc - shaped sacs or cisternae .
- Golgi complex is made up of four parts : cisternae , tubules , vesicles and Golgian vacuoles .
- One face of Golgi apparatus is convex or forming ( cis face ) while other is concave side or maturing face ( trans face ) .
- Golgi apparatus helps in processing , packaging , transport and release of secretory proteins .
- Glycosylation of proteins and lipids to form glycoproteins and glycolipids .
- They act as primary lysosomes as they store digestive enzymes obtained through ER in the inactive state .
- Transformation and recycling of plasma membrane takes place here .
- These were first reported by Christian de Duve in 1955 through fractionation technique .
- They are found in all animal cells except RBCs . In plants and fungi , their function is taken over by vacuoles.
- These are single membrane bound , small vesicular organelles containing hydrolytic enzymes.
- The organelle passes through various stages and shows polymorphism .
- There are four types of lysosomes :
- ( i ) primary lysosome , ( ii ) secondary lysosome , ( iii ) residual bodies and ( iv ) autophagic vacuoles .
- They help in digestion of food obtained through phagocytosis .
- Harmful and unwanted materials are disposed off by lysosomes . Thus , lysosomes are called disposal bags or units .
- In injured and dead cells , the lysosome membrane ruptures spontaneously releasing the digestive enzymes or acid hydrolases that lyse the weakened cells. Therefore , these are called ' suicide bags ' of the cell.
Vacuoles
- Vacuoles are non - cytoplasmic areas present inside the cytoplasm .
- These are surrounded by single membrane called tonoplast .
- These are formed by expansion and pinching off from endoplasmic reticulum .
- They are small in animal cells and large in fungal and plant cells .
- Depending upon the contents and function , these are of four types - sap vacuoles , food vacuoles , contractile vacuoles and air vacuoles .
- Vacuoles maintain osmotic pressure and turgidity and also help in osmoregulation . They provide buoyancy , mechanical strength and protection .
- In plants, the tonoplast facilitates the transport of a number of ions and other materials against concentration gradient .
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